Grizzly 399, a beloved mama bear known as the queen of Wyoming's Grand Teton National Park, has died after being hit by a car, the U.S. Fish & Wildlife Service reports.

Grizzly 399 is Mangelsen’s muse. “She’s beautiful in ways we think the classic grizzly bear might be described,” Mangelsen says. “Her personality is calm, generally speaking.

Grizzly No. 399 and her ever-growing brood was a beloved favorite of tourists visiting Wyoming’s Grand Teton National Park for nearly 30 years before her untimely death on Highway 29/28 on Oct. 22.

Grizzly 399's death marks the second bear mortality this year caused from a vehicle strike in the Greater Yellowstone Ecosystem, according to the U.S. Fish & Wildlife Service.

Jacksonites may remember when Grizzly 399, the fabled Queen of the Tetons, was caught on camera strolling with her cubs past ...

Ashes remains of Grizzly Bear No. 399 returned to Pilgrim Creek area Grand Teton National Park Yellowstone Wyoming after she was killed by vehicle 18 known cubs.

Grizzly No. 399 continued to reproduce in old age and raised several cubs close to Grand Teton National Park's roads. As a result, visitors eagerly watched her in her natural habitat, per CNN.

“The bear’s remains were put back onto the landscape in Grand Teton National Park.” Grizzly 1058’s mother was Grizzly 399, a well-known bear that also lived in Grand Teton National Park.

In this undated photo provided by Grand Teton National Park a grizzly bear known as No. 399 stands along side a cub. (C. Adams/Grand Teton National Park via AP) \CHEYENNE, Wyo. (AP) — A famous grizzly ...

Grizzly 399 has come out with four cubs and that, in itself, is quite unheard of. Not too long ago, the bears were declining so fast that it was believed they would all be extinct by now.

Grizzly 399, a beloved mama bear known as the queen of Wyoming's Grand Teton National Park, has died after being hit by a car, the U.S. Fish & Wildlife Service reports.

Grizzly bear #399 facts: 28 years old; one of only 12 known grizzly bears in the Greater Yellowstone Ecosystem to reach the age of 28 or older, representing 0.6% of all bears documented and aged ...