Grizzly 399, a beloved mama bear known as the queen of Wyoming's Grand Teton National Park, has died after being hit by a car, the U.S. Fish & Wildlife Service reports.

By clicking Sign up, you agree to our privacy policy. Named for the tag affixed by researchers to her ear, No. 399 was the oldest-known reproducing female grizzly in the greater Yellowstone ecosystem.

Grizzly No. 399 and her ever-growing brood was a beloved favorite of tourists visiting Wyoming’s Grand Teton National Park for nearly 30 years before her untimely death on Highway 29/28 on Oct. 22.

“The bear’s remains were put back onto the landscape in Grand Teton National Park.” Grizzly 1058’s mother was Grizzly 399, a well-known bear that also lived in Grand Teton National Park.

Grizzly No. 399 continued to reproduce in old age and raised several cubs close to Grand Teton National Park's roads. As a result, visitors eagerly watched her in her natural habitat, per CNN.

Jacksonites may remember when Grizzly 399, the fabled Queen of the Tetons, was caught on camera strolling with her cubs past ...

Grizzly bear No. 399 and her four cubs cross a road as Cindy Campbell stops traffic in Jackson Hole, Wyo., on Nov. 17, 2020. Credit: AP/Ryan Dorgan ...

JACKSON, Wyo. — “There has never been a bear in history that has beaten the odds like 399,” her quasi-official photographer, Thomas D. Mangelsen, said of the world-famous grizzly in February.

Grizzly 399, she learned over time that if she hangs out within 100 yards of the road, she has these people around that are in a way protecting her cubs from the male grizzly bears, which is very ...

Jacob Krank, the master of ceremonies, shared how he encountered No. 399 the first time he drove into the park to explore it 13 years ago. Suddenly he saw “this beautiful grizzly bear” in the ...

Credit: AP A shrine sits on the corner of town square to mourn the loss of grizzly bear No. 399 at a candlelight vigil in Jackson, Wyo. Author: Associated Press Published: 2:46 PM EST November 3 ...